It is now generally accepted that the roots of our ethics lie in patterns of behavior that evolved among our pre-human ancestors, the social mammals and that we retain within our biological nature elements of these evolved responses. We have learned considerably more about this responses, and we are beginning to to understand how they interact with our capacity to reason.
Preface To The 2011 edition, p. xi - The Expanding Circle: Ethics, Evolution, and Moral Progress (1981)