The theory of the origin of new species by variation will also explain why a species which has once died out never reappears, and why the fossil fauna and flora recede farther and farther from the living type in proportion as we trace it back to remoter ages. It would also account for the fact, that when we have to intercalate a new set of fossiliferous strata between two groups previously known, the newly discovered fossils serve to fill up gaps between specific or generic types previously familiar to us, supplying often the missing links of the chain, which, if transmutation is accepted, must once have been continuous.

Ch.21, p. 414-415 - The Geological Evidences of the Antiquity of Man (1863)